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The pace of infilling of the lagoon has increased over the last 6,000 years, along with accelerated growth of sandspits and the creation of new inland marshes and wetlands.The extensive marshes of Doñana National Park now have a flat topography, with some inland depressions occupied by temporary or permanent wetlands, locally called 'lucios'.

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The eco-system has been under constant threat by the draining of the marshes, the use of river water to boost agricultural production by irrigating land along the coast, water pollution by upriver mining, and the expansion of tourist facilities. Doñana National Park has a biodiversity that is unique in Europe, although there are some similarities to the Parc Naturel Régional de Camargue of the Camargue river delta in France, with which Doñana Park is twinned.

The park features a great variety of ecosystems and shelters wildlife including thousands of European and African migratory birds, fallow deer, Spanish red deer, wild boars, European badgers, Egyptian mongooses, and endangered species such as the Spanish imperial eagle and the Iberian lynx.

Reference to the use of Coto Donana as a hunting lodge is made in the first verses of the La Fábula de Polifemo y Galatea (Fable of Polyphemus and Galatea), which the lyric poet Luis de Góngora dedicated to the Count of Niebla, and in which he requests that the nobles suspend their hunting exploits to hear his verses.

In 1624, King Philip IV stayed at the estate for several days as a guest of the 9th Duke of Medina Sidonia, and joined in some large hunts.

The geological profile of Doñana National Park reflects the development over several hundred thousand years of a deep aquifer and geomorphological features that have enhanced the biodiversity of the wildlife habitats presently found there.

After the end of the last glacial period, the area was covered by freshwater and brackish marshes, ponds and sand dunes, with some marine intrusions caused by high-energy events such as tsunamis and large storms.

The park is an area of marshes, shallow streams, and sand dunes in Las Marismas, the delta where the Guadalquivir River flows into the Atlantic Ocean.

It was established as a nature reserve in 1969 when the World Wildlife Fund joined with the Spanish government and purchased a section of marshes to protect it.

He brought with him a great variety of comestibles, which included such luxuries as snow from the mountains for the frequent banquets he held in honor of the duke.

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